
His ribs spread out to act as a border for the gate, and then double as large extensible teeth that can consume everything in his eye's field of vision in an instant. When activating this portal, Gluttony's stomach opens up, revealing a large eye at its center.

In the manga, Gluttony is the failed product of Father 's attempts to create a Portal of Truth.

He particularly likes eating people, and the only way he can find enjoyment in a battle is if he can eat his opponent afterwards He has powerful jaws and acidic saliva, and can and will eat almost anything. He is a large, simple-minded homunculus whose thoughts rarely stray far from eating. Gluttony ( グラト二ー, Guratonī), the second homunculus introduced in the series, typically appears in the company of Lust. Kikuko Inoue voices her in Japanese in the new series. Yūko Satō voices her in Japanese, and Laura Bailey in English. Wrath eventually catches up to them before he paralyzes her using a locket filled with the hair from her original form and kills her. This ultimately leads to her defection from the homunculi, and to her aiding Edward Elric if he promises to help make her human.
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Though originally largely the same as her manga counterpart, Lust, as the series progresses, begins to develop a strong desire to regain her humanity. In the first anime, Lust was created when Scar's brother tried to revive his deceased lover. Mustang repeatedly blasts Lust with flames, ultimately killing her after depleting the power of her Philosopher's Stone. In the manga, after leading an effort to capture Barry the Chopper, her plans backfire when Roy Mustang infiltrates the homunculi's secret lair. She can extend her fingers to great lengths, which are capable of cutting through virtually any substance. Lust Lust ( ラスト, Rasuto), the "Ultimate Spear," appears as the first homunculus in the series as a shapely woman who primarily works in the shadows, acts as envoy for her leader in both iterations, and encourages alchemists down her desired path. The homunculi of the first anime possess an additional weakness: a remnant of their original bodies (a bone, hair, etc.), which weaken and immobilize them when they come in contact with them. These homunculi are led by Dante, and feed on incomplete Philosopher's Stones to fuel their powers they are susceptible to death once these stones are regurgitated. In the first anime, homunculi are the result of a human transmutation, and closely resemble the deceased human intended to be revived. The only way to kill a homunculus is by expending all of the power in their Philosopher's Stone, leaving them unable to revive themselves once they have been killed. Others, such as Wrath, are humans that accepted a Philosopher's Stone into their body, gaining the abilities of a homunculus while aging like a human. In the manga, the homunculi were created by or are at the discretion of Father, being a manifestation of one of his traits: his greed, his lust, and so onEach homunculus has a Philosopher's Stone somewhere in his or her body which fuels their regenerative ability, and were created at the same time as their respective Philosopher's Stone. Aside from this, the concept of the homunculi varies drastically between the manga and first anime series.


They also possess a heightened regenerative ability, rendering them nigh indestructible, though not invincible. Each homunculus is named after one of the seven deadly sins, and can be identified by their bearing of the mark of the Ouroboros somewhere on their body. They are a group of artificially created humans who seek to fulfill their leader's goals of creating a Philosopher's Stone. From left to right: Lust, Envy, Sloth, Gluttony, Pride, Wrath, and Greed in the manga Homunculi ( ホムンクルス, Homunkurusu) serve as the main antagonists in Fullmetal Alchemist.
